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[Kumis]

According to what I've read, If you've found a cavern in a previous fort then you can have moss grow in another fort because the cavern connects the two forts. Unfortunately you can't travel to the old fort in Dwarf Mode.

Moss will always grow once you uncovered it in fortress mode. There seems to be one flag "have moss" per world.

I don't know if I believe that. Unless I remember incorrectly my other (intermediate) forts in the same world didn't have moss grow.
It's only this one, and so it's probably either the same civilisation, the same cavern system or both.

Do you have a source?

[gestahl]

It's from having breached a cave in a different fort. I've got no idea if being the same civ matters. I do know that it is not a world-wide effect, but is bigger than just one biome. I'd suspect it's either per-region or per-16x16 embark squares.

[Exitstrategy]

"Once an underground cavern has been discovered, shrubs and trees will spontaneously grow on any subterranean soil or muddied rock on every embark site that accesses the cavern. This means that if you find a cavern in one embark site and embark in another site accessing the same cavern, plants will start growing there even before you discover the branch of the cavern that lies under the site. This allows you to construct underground tree farms and avoid sending dwarves to the surface to harvest wood, or just to get wood in environments without above-ground forests."

-Taken from Benefits section of the wiki page here

[Kumis]

That'd be it then. Cheers!

[SixOfSpades]

Theoretically, it's even possible to have underground flora & knowledge of magma furnaces in the first fort you start in a world, and well before breaching the caverns. This is because some caverns are generated with caves already connecting them with the surface, so they're growing floor fungus from worldgen on. And if you embark atop a cavern that has one of these caves (even if the cave itself is not on your embark), then that cavern is considered "breached," and you get cave trees.